Rod Blackshaw

Professor Rod Blackshaw

Job title: Professor of Agricultural Zoology, School of Biomedical and Biological Sciences (Faculty of Science and Technology)
Address: B426, Portland Square, Drake Circus,
Plymouth, Devon, PL4 8AA
Email: R.Blackshaw@plymouth.ac.uk

Role
Director, Centre for Agricultural and Rural Sustainability

Qualifications & background

BSc in Agricultural Zoology University of Newcastle upon Tyne

PhD in ecoxicology and soil fauna, University of Newcastle upon Tyne

Employed from 1981 to 1996 as a research scientist, Departmentof Agriculture for Northern Ireland

Moved to the University of Plymouth in 1996 as Head of the Department of Agriculture and Food Studies.

Appointed Dean of the Faculty of Land, Food & Leisure in 2001.

Transfered to the School of Biological Sciences in 2003.

Professional membership

Fellow of the Royal Entomological Society

Board Member, Soil Ecology Society

Roles on external bodies

Current activities:
1. Member of Sustainable Arable LINK Programme Management Committee (1998 - ). Advises DEFRA, SERAD and BBSRC on research funding allocation. Monitor progress of projects.

2. Proposal & programme reviewer for DEFRA.

3. Proposal Reviewer, BBSRC

4. Proposal Reviewer Technology Strategy Board

6. Project evaluator ESRC (RELU)

7. Proposal Reviewer Norwegian Research Council

8. Referee for various journals

Past activities:
1.Member, Research and Education Advisory Panel Department of Agriculture and Rural Development for Northern Irealnd (2006 - 2009 ).

2. Invited Reviewer, DEFRA Arable Zoology and Pest Management Programme (2004).

3. Visiting Group Member, DEFRA Central Sciences Laboratory (2001)

4. Invited Contributor, CABI Pest Compendium CD-Rom Database (2000)

5. Invited Member MAFF Plant Health Research Review Group (1999)

6. Invited Reviewer, Scottish Office Plant Sciences Research Programme (1999)

7. Council member of Royal Entomological Society (1996-1999; 2008-2011)

Teaching interests

Integrated pest management, invertebrate ecology.

Staff serving as external examiners
External Examiner, Environmental Science, Nottingham Trent University

Research interests

Research activities fit under two broad headings:

INTEGRATED PEST MANAGEMENT The majority of pest managment practices have hardly evolved since the concept of Integrated Control first emerged in the 1960s. They remain centred on managment practices directed against target pests in crops, using the field as the key managment unit. My research is directed towards unravelling the broader ecology of pest species, understanding how they function within agricultural landscapes, and identifying new intervention points to reduce damaging population levels. Much of this research has been directed towards the spatio-temporal behaviours and distributions of organisms, but more recent work has revealed that although monitoring is a recognised key activity in pest management, direct interpretation of trap counts is challenging because we have no means of separating density and activity components of counts. Thus much of my current activity is directed towards understanding the behaviour of individuals around traps, and the interpretation of this using mathematical models, working with Dr Sergei Petrovskii at Leicester University.

SOIL BIOLOGY In collaboration with Dr Phil Murray of Rothamsted, North Wyke we have developed new methods of tracking the flow of carbon and nitrogen through soil food webs. This work has shown that in soil food webs omnivorous animals may be functionally selective so that they can occur in different trophic levels, and that previously assumed functional/trophic ideas underestimate the complexity. This remains an on-going research interest. Spatial ecology also plays a part in understanding how soils operate as biological systems. Research into this aspect is being addressed through the development of novel sampling methods and the development of theoretical understanding of how complex physical environments affect animal movement behaviours.

The current team comprises 1 PDRA, and 4 PhD and 1 ResM students:

1. Susan Morris - dynamics of rare arable weeds (with Dr Miguel Franco).
2. Ayano Saski-Crawley - behaviour of potato cyst nematodes and potential trap crops (with Prof John Pickett, Rothamsted Harpenden)
3. Jenny Williams - dispersal of soil mesofauna (with Dr Phil Murray, Rothamsted North Wyke)
4. Nawzad kadir - effects of soil biodiversity on plant-herbivore interactions (with Dr Mick Hanley)
5. Karzan D. Ahmed - spatial distribution of soil insects (with Dr Phil Murray, Rothamsted North Wyke)

6. Dr Carly Benefer - Theoretical and experimental understanding of insect trapping systems (with Dr Sergei Petrovskii, University of leicester).

Active external collaborators are Dr Bob Vernon (Agriculture and AgriFood Canada), Dr Sergei Petrovskii (University of Leicester), Dr Dan Peck (Corvallis University) .

UoP Research group membership

Centre for Agricultural and Rural Sustainability (CARS)
Terrestrial Ecology Research Group

Other research

Conferences organised
Soil Entomology and Ecology, Plymouth 2011

7th International Colloquium on Earthworm Ecology, Cardiff, 2002.

Entomology 2008 for the Royal Entomological Society

Entomology 2011 Crop Protection Session Organiser

Chapters

Zuur, A F, ., Blackshaw, RP., Vernon, R S, ., Thiebaud, F, . & Ieno, E N, . (2012) 'Analysis of zero inflated click-beetle data' in Zurr AF,Saveliev, A A ,Ieno, E N (ed.) Zero inflated models and generalized mixed models with R Newburgh, UK: Highland Statistics Ltd pp 257 - 276

Blackshaw, RP. (2009) 'Predicting the size of leatherjacket populations in grassland: a pilot study using water traps' in Ehlers R-U,Crickmore N,Enkerli J,Glazer I,Lopez-Ferber M,Tkaczuk C (ed.) Working Group on Insect Pathogens and Insect Parasitic Nematodes and COST Action 862 Bacterial Toxins for Insect control. 12th Meeting Future Research and Development in the Use of Microbial Agents and Nematodes for Biological Insect Control. Proceedings of the meeting at Pamplona, Spain, 22nd to 25th June 2009 International Organisation for Biological Control pp 426 - 429 http://www.iobc-wprs.org/pub/bulletins/bulletin_2009_45_table_of_contents_abstracts.pdf

Blackshaw, RP. & Kerry, BJ. (2009) 'Root herbivory in arable systems' in Johnson SN,Murray PJ (ed.) Root feeders: an ecosystem perspective CABI Publishing pp 33 - 53

Blackshaw, RP., Hicks, H. & Vernon, RS. (2009) 'Sex pheromone traps for predicting wireworm populations: limitations to interpretation' in Collier R (ed.) Working Group on Integrated Protection of Field Vegetables: Proceedings of the Meeting at Porto, Portugal, 23rd to 29th September 2007 International Organisation for Biological Control pp 17 - 21 http://www.iobc-wprs.org/pub/bulletins/bulletin_2009_51_table_of_contents_abstracts.pdf

Benefer, C., Blackshaw, RP., Knight, M. & Ellis, J. (2009) 'The relationship between Agriotes click beetle (Coleoptera: Elateridae), wireworms and environmental factors' in Ehlers R-U,Crickmore N,Enkerli J,Glazer I,Lopez-Ferber M,Tkaczuk C (ed.) Working Group on Insect Pathogens and Insect Parasitic Nematodes and COST Action 862 Bacterial Toxins for Insect control. 12th Meeting Future Research and Development in the Use of Microbial Agents and Nematodes for Biological Insect Control. Proceedings of the meeting at Pamplona, Spain, 22nd to 25th June2009 International Organisation for Biological Control p 425 http://www.iobc-wprs.org/pub/bulletins/bulletin_2009_45_table_of_contents_abstracts.pdf

Blackshaw, RP. (2008) 'When should we use biocontrol agents against leatherjackets (Tipula paludosa Meig.)?' Working Group on Insect Pathogens and Insect Parasitic Nematodes: Proceedings of the Meeting at Alés, France, 3rd to 7th June 2007 International Organisation for Biological Control pp 313 - 318 http://www.iobc-wprs.org/pub/bulletins/bulletin_2008_31_table_of_contents_abstracts.pdf

Blackshaw, RP., Vernon, RS. & Hicks, H. (2007) 'Spatial distribution of click beetles (Agriotes spp.) at field and landscape scales' Working Group on Insect Pathogens and Insect Parasitic Nematodes, Subgroup on Soil Insect Pests: Proceedings of the meeting at Auer/Ora, Italy, 16th to 18th October 2006 International Organisation for Biological Control pp 27 - 33 http://www.iobc-wprs.org/pub/bulletins/bulletin_2007_30_07_abstracts.pdf

Blackshaw, RP. & Petrovskii, SV. (2007) 'Tipula paludosa population dynamics: challenginng the myth of environmental limitation' in Enkerli J (ed.) Working Group on Insect Pathogens and Insect Parasitic Nematodes, Subgroup on Soil Insect Pests: Proceedings of the meeting at Auer/Ora, Italy, 16th to 18th October 2006 International Organisation for Biological Control pp 115 - 118 http://www.iobc-wprs.org/pub/bulletins/bulletin_2007_30_07_abstracts.pdf

Thomas, CFG., Blackshaw, RP., Hutchings, L., Woolley, C., Goodacre, S., Hewitt, GM., Ibrahim, K., Brooks, SP. & Harrington, R. (2003) 'Modelling life-history/dispersal strategy interactions to predict and manage linyphid spider diversity in agricultural landscapes' in Rossing WAH,Poehling H-M,Burgio G (ed.) Landscape Management for Functional Biodiversity: Proceedings of 1st Meeting in Bologna, Italy, 11th to 14th May 2003 International Organisation for Biological Control pp 167 - 172

Blackshaw, RP., Winder, L. & Lefley, M. (1998) 'Potential of fuzzy logic in crop protection decision making' in British Crop Protection Council (ed.) The 1998 Brighton Conference: Pests & Diseases: Proceedings of an International Conference, Brighton, November 1998 [3 volumes] British Crop Protection Council pp 446 - 454

Blackshaw, RP. (1996) 'Importance of overwintering adults to summer oviposition in Northern Ireland' in Alford DV,Backhaus GF (ed.) Second International Workshop on Vine Weevil (Otiorhynchus sulcatus Fabr.) (Coleoptera: Curculionidae): Brunswick, Germany, 21st - 23rd May 1996 Blackwell Wissenschafts-Verlag pp 36 - 40

McCracken, AR. & Blackshaw, RP. (1993) 'The protection of the Northern Ireland nursery stock industry from harmful organisms' in Kavanagh JA,Brennan P,Academy RI (ed.) Plant Health and 1992: Proceedings of a seminar held at the Royal Irish Academy, Dublin on 8th and 9th March 1990 Dublin: Royal Irish Academy pp 55 - 66

Blackshaw, RP. & Diggle, PJ. (1985) 'Effect of some chemicals on earthworm populations' in Andrews DF,Herzberg AM (ed.) Data: a collection of problems from many fields for the student and research worker New York: Springer Verlag pp 201 - 206

Conference Papers

Benefer, C., Blackshaw, R., Knight, M. & Ellis, J. (2011) 'Development of amplified fragment length polymorphism (AFLP) markers for assessing Agriotes click beetle dispersal in agricultural land' in Ehlers R,Crickmore N,Enkerli J,Glazer I,Kirchmair M,Lopez-Ferber M,Neuhauser S,Strasser H,Tkaczuk C,Traugott M (ed.) Insect Pathogens and Entomopathogenic Nematodes Biological Control in IPM Systems. Innsbruck, Austria 19/06/2011 - 23/06/2011 Darmstadt, Germany: International Organization for Biological and Integrated Control of Noxious Animals and Plants, West Palearctic Regional Section (IOBC/WPRS) pp 541 - 541 http://www.iobc-wprs.org/

Benefer, C., Blackshaw, RP., Knight, ME. & Ellis, JS. (2011) 'Male Agriotes click beetle walking behaviour and response to sex pheromones' in Ehlers RU,Crickmore N,Enkerli J,Glazer I,Kirchmair M,Lopez-Ferber M,Neuhauser S,Strasser H,Tkaczuk C,Traugott M (ed.) Insect Pathogens and Entomopathogenic Nematodes Biological Control in IPM Systems. Innsbruck, Austria 19/06/2011 - 23/06/2011 Darmstadt, Germany: International Organisation for Biological and INtegrated Control of Noxious Animals and Plants, West Palearctic Regional Section (IOBC/WPRS) pp 504 - 504 http://www.iobc-wprs.org/

Benefer, C., Blackshaw, RP., Knight, ME. & Ellis, JS. (2009) 'The relationship betweenAgriotes click beetle (Coleoptera: Elateridae) wireworms and environmental factors' in Ehlers RU,Crickmore N,Enkerli J,Glazer I,Lopez-Ferber M,Tkaczuk C (ed.) Insect Pathogens and Insect Parasitic Nematodes. Pamplona, Spain 22/06/2009 - 25/06/2009 International Organisation for Biological and Integrated Control of Noxious Animals and Plants, West Palearctic Regional Section (IOBC/WPRS) p 425 http://www.iobc-wprs.org/

Blackshaw, RP., Benefer, C., Knight, MK., Ellis, J., Hicks, H. & Vernon, RS. (2009) 'Recent advances in wireworm research: new approaches raise new questions' Crop Protection in Southern Britain: an Association of Applied Biologists Conference. East of England Showground, Peterborough 10/02/2009 - 11/02/2009 pp 99 - 103

Blackshaw, RP. (2009) 'INTRA-GENERATIONAL MORTALITY OF LEATHERJACKETS (TIPULA PALUDOSA).' SOC NEMATOLOGISTS pp 310 - 311

Crotty, FV., Adl, SM., Clegg, CD., Blackshaw, RP. & Murray, PJ. (2009) 'INVESTIGATING SOIL FOOD WEBS - TRACKING THE TRANSLOCATION OF C-13 AND N-15 THROUGH MICROBIAL INTERACTIONS WITHIN THE SOIL.' SOC NEMATOLOGISTS pp 321 - 322

Blackshaw, RP. (2009) 'PREDICTING THE SIZE OF LEATHERJACKET (TIPULA PALUDOSA MEIG.) POPULATIONS IN GRASSLAND.' SOC NEMATOLOGISTS pp 311 - 311

Benefer, C., Blackshaw, R., Knight, M. & Ellis, J. (2009) 'THE DISTRIBUTION OF SOIL INSECTS OVER DIFFERENT SPATIAL SCALES IN GRASSLAND.' SOC NEMATOLOGISTS pp 309 - 309

Blackshaw, RP., Hicks, H. & Vernon, RS. (2008) 'Sex pheromone traps for monitoring wireworm populations: how effective are they?' Diversifying Crop Protection: Endure International Conference 2008. La Grande Motte, France 12/10/2008 - 15/10/2008 Endure Network p 0.39 http://www.endure-network.eu/international_conference_2008/proceedings/wednesday_october_15

Blackshaw, RP. (2008) 'Using population models to develop management tactics for Tipula paludosa in organic systems' Diversifying Crop Protection: Endure International Conference 2008. La Grande Motte, France 12/10/2008 - 15/10/2008 Endure Network p O.13 http://www.endure-network.eu/international_conference_2008/proceedings/tuesday_october_14

Blackshaw, RP., Donovan, SE., Hazarika, S., Bol, R. & Dixon, ER. (2007) 'Earthworm responses to long term agricultural management practices: Spatial relationships with soil properties' 8th International Symposium on Earthworm Ecology (ISEE8). Cracow, POLAND 04/09/2006 - 09/09/2006 ELSEVIER FRANCE-EDITIONS SCIENTIFIQUES MEDICALES ELSEVIER pp S171 - S175 , DOI: 10.1016/j.ejsobi.2007.08.014

Petrovskii, SV. & Blackshaw, RP. (2005) 'Gone with the wind: is it always true for invasive plants and, if not, why not?' in Alford DV,Backhaus GF (ed.) Plant protection and Plant Health in Europe: Introduction and Spread of Invasive Species. Alton, Hampshire: BCPC: British Crop Production Council pp 277 - 278

Blackshaw, RP. (1999) 'Behaviour mediated population regulation?' Challenges in Applied Population Biology: an Association of Applied Biologists Conference. London pp 125 - 130

Blackshaw, RP. (1997) 'Life cycle of the earthworm predator Artioposthia triangulata (Dendy) in northern Ireland' 5th International Symposium on Earthworm Ecology (ISEE 5). COLUMBUS, OH 05/07/1994 - 09/07/1994 PERGAMON-ELSEVIER SCIENCE LTD pp 245 - 249

Blackshaw, RP. (1997) 'The planarian Artioposthia triangulata (Dendy) feeding on earthworms in soil columns' 5th International Symposium on Earthworm Ecology (ISEE 5). COLUMBUS, OH 05/07/1994 - 09/07/1994 PERGAMON-ELSEVIER SCIENCE LTD pp 299 - 302

Blackshaw, RP. (1996) 'Control options for the New Zealand flatworm' Brighton Crop Protection Conference on Pests and Diseases - 1996. BRIGHTON, ENGLAND 18/11/1996 - 21/11/1996 BRITISH CROP PROTECTION COUNCIL pp 1089 - 1094

Coll, C. & Blackshaw, RP. (1996) 'Leatherjackets in winter cereals: a self-inflicted problem?' Rotations and Cropping Systems: an Association of Applied Biologists Conference. Cambridge, UK pp 145 - 151

Blackshaw, RP. (1995) 'Effect of tank-mixing on economic thresholds for pest control' Understanding Crop Protection Mixtures: an Association of Applied Biologists Conference. Silsoe 11/04/1995 - 12/04/1995 pp 41 - 49

Blackshaw, RP., Stewart, RM., Humphreys, IC. & Coll, C. (1994) 'Preventing leatherjacket damage in cereals' Sampling to Make Decisions: an Association of Applied Biologists Conference. Cambridge, UK 22/03/1994 - 23/03/1994 pp 189 - 196

Blackshaw, RP. (1994) 'Sampling leatherjackets in grassland' Sampling to Make Decisions: an Association of Applied Biologists Conference. Cambridge, UK 22/03/1994 - 23/03/1994 pp 95 - 102

Coll, C., Blackshaw, RP., Humphreys, I. & Stewart, RM. (1993) 'The epidemiology of Tipula oleracea in winter cereals in Northern Britain' Troisieme Conference Internationale sur les Ravageurs en Agriculture [3rd International Conference on Pests in Agriculture]. Montpellier 07/12/1993 - 09/12/1993 Paris: Association Nationale de Protection des Plantes (ANPP) pp 1059 - 1066

Blackshaw, RP., Humphreys, IC., Coll, C. & Stewart, RM. (1993) 'A nnew leatherjacket pest configuration for Northern Ireland' Crop Protection in Northern Britain 1993. University of Dundee Scottish Crop Research Institute p 365

Blackshaw, RP. (1993) 'Current status of the New Zealand flatworm' Crop Protection in Northern Britain 1993. University of Dundee Scottish Crop Research Institute p 367

Humphreys, IC., Blackshaw, RP., Stewart, RM. & Coll, C. (1993) 'Distribution of Tipula oleracea and Tipula paludosa in Northern Britain' Crop Protection in Northern Britain 1993. University of Dundee Scottish Crop Research Institute pp 85 - 90

Stewart, RM., Humphreys, IC., Blackshaw, RP. & Coll, C. (1993) 'Sampling techniques for leatherjackets in arable situations' Crop Protection in Northern Britain 1993. University of Dundee Scottish Crop Research Institute p 364

Coll, C., Greenhorn, JG. & Blackshaw, RP. (1993) 'The influence of oilseed rape (Brassica napus) on the occurrence of leatherjackets (Tipula spp.) in winter cereals' Crop Protection in Northern Britain 1993. University of Dundee Scottish Crop Research Innstitute pp 79 - 84

Mowat, DJ. & Blackshaw, RP. (1991) 'Can established grass pests be controlled by one spray?' Strategies for Weed, Disease and Pest Control in Grassland. Gloucester British Grassland Society pp 10.17 - 10.18

Blackshaw, RP. (1991) 'Leatherjackets in grassland' Strategies for Weed, Disease and Pest Control in Grassland. Gloucester British Grassland Society pp 6.1 - 6.12

Blackshaw, RP. (1990) 'Forecasting leatherjacket populations in Northern Ireland' Crop Protection in Northern Britain 1990. University of Dundee Scottish Crop Research Institute pp 117 - 122

Kell, GV. & Blackshaw, RP. (1987) 'Grassland characteristics influencing leatherjacket populations' Crop Protection in Northern Britain 1987. University of Dundee 15/03/1987 - 17/03/1987 Scottish Crop Research Institute p 422

Blackshaw, RP. (1987) 'Status of leatherjackets as grassland pests in Northern Ireland' Crop Protection in Northern Britain 1987. University of Dundee 15/03/1987 - 17/03/1987 Scottish Crop Research Institute p 422

Kelly, MS. & Blackshaw, RP. (1987) 'Studies on natural enemies of leatherjacket populations in Northern Ireland' Crop Protection in Northern Britain 1987. University of Dundee 15/03/1987 - 17/03/1987 Scottish Crop Research Institute p 423

D'Arcy-Burt, S. & Blackshaw, RP. (1986) 'Distribution studies on grassland Bibionidae' Dipterology: 1st International Congress. Budapest Budapest: Hungarian Academy of Sciences Biological Section p 52

D'Arcy-Burt, S. & Blackshaw, RP. (1985) 'A distribution study of bibionid larvae in grassland' in Brockman JS (ed.) Weeds, Pests and Diseases of Grassland and Herbage Legumes. University of Nottingham 10/04/1985 - 12/04/1985 Croydon: British Crop Protection Council p 301

Blackshaw, RP. (1984) 'Frequency of spring barley damage by leatherjackets in Northern Ireland' British Crop Protection Conference: Pests and Diseases. Croydon: British Crop Protection Council pp 15 - 18

Blackshaw, RP. (1984) 'Studies on the chemical control of vine weevil larvae' British Crop Protection Conference: Pests and Diseases. Croydon: British Crop Protection Council pp 1093 - 1098

Kell, GV. & Blackshaw, RP. 'The influence of agricultural practices on leatherjacket populations in Northern Ireland' Grassland and Animal Production, Now and in the Future. Dublin European Grassland Federation pp 504 - 508

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Journals

Articles

Crotty, FV., Adl, SM., Blackshaw, RP. & Murray, PJ. (2012) 'Using Stable Isotopes to Differentiate Trophic Feeding Channels within Soil Food Webs.' J Eukaryot Microbiol , , DOI: 10.1111/j.1550-7408.2011.00608.x

Crotty, FV., Blackshaw, RP. & Murray, PJ. (2011) 'Differential growth of the fungus Absidia cylindrospora on 13C/15N-labelled media.' Rapid Commun Mass Spectrom 25 (11), pp 1479 - 1484 , DOI: 10.1002/rcm.4888

Crotty, FV., Adl, SM., Blackshaw, RP. & Murray, PJ. (2011) 'ESTIMATING N AND C TRANSFORMATIONS FROM STABLE ISOTOPE TRACER STUDIES IN AN INTACT SOIL FOOD WEB' J PHYCOL 47, pp S7 - S7

Crotty, FV., Blackshaw, RP. & Murray, PJ. (2011) 'Tracking the flow of bacterially derived 13C and 15N through soil faunal feeding channels.' Rapid Commun Mass Spectrom 25 (11), pp 1503 - 1513 , DOI: 10.1002/rcm.4945

Benefer, C., Andrew, P., Blackshaw, R., Ellis, J. & Knight, M. (2010) 'The spatial distribution of phytophagous insect larvae in grassland soils' APPL SOIL ECOL 45 (3), pp 269 - 274 , DOI: 10.1016/j.apsoil.2010.05.002

Blackshaw, RP. (2009) 'A comparison of management options for leatherjacket populations in organic crop rotations using mathematical models' AGR FOREST ENTOMOL 11 (2), pp 197 - 203 , DOI: 10.1111/j.1461-9563.2008.00426.x

Murray, PJ., Clegg, CD., Crotty, FV., Martinez, ND., Williams, JK. & Blackshaw, RP. (2009) 'Dissipation of bacterially derived C and N through the meso- and macrofauna of a grassland soil' SOIL BIOL BIOCHEM 41 (6), pp 1146 - 1150 , DOI: 10.1016/j.soilbio.2009.02.021

Ellis, JS., Blackshaw, R., Parker, W., Hicks, H. & Knight, ME. (2009) 'Genetic identification of morphologically cryptic agricultural pests' AGR FOREST ENTOMOL 11 (1), pp 115 - 121 , DOI: 10.1111/j.1461-9563.2008.00420.x

Crotty, FV., Clegg, CD., Blackshaw, RP. & Murray, PJ. (2009) 'Unearthing links within the soil food web: following microbial carbon and nitrogen flow through the trophic levels' Antenna (Bulletin of the Royal Entomological Society) 33 (2), pp 149 - 150

Blackshaw, RP. (2009) 'Wireworms. Problems with sex pheromones and uncertainty' TAG Bulletin 22, pp 12 - 13

Petrovskii, SV., Blackshaw, RP. & Bai-Lain, L. (2008) 'Consequences of the allee effect and intraspecific competition on population persistence under adverse environmental conditions' Bulletin of Mathematical Biology 70 (2), pp 412 - 437 http://www.springerlink.com/content/0092-8240

Hicks, H. & Blackshaw, RP. (2008) 'Differential responses of three Agriotes click beetle species to pheromone traps' AGR FOREST ENTOMOL 10 (4), pp 443 - 448 , DOI: 10.1111/j.1461-9563.2008.00397.x

Blackshaw, RP. & Vernon, RS. (2008) 'Spatial relationships between two Agriotes click-beetle species and wireworms in agricultural fields' Agricultural and Forest Entomology 10 (1), pp 1 - 11 http://onlinelibrary.wiley.com/doi/10.1111/afe.2008.10.issue-1/issuetoc , DOI: 10.1111/j.1461-9563.2007.00351.x

Blackshaw, RP., Vernon, RS. & Hicks, H. (2008) 'The wireworm pest complex: evidence for species differences from spatial studies.' Commun Agric Appl Biol Sci 73 (3), pp 429 - 432

Blackshaw, RP. & Petrovskii, SV. (2007) 'Limitation and regulation of ecological populations: a meta-analysis of Tipula paludosa field data.' Mathematical Modelling of Natural Phenomena 2 (4), pp 46 - 62

Blackshaw, RP. & Vernon, RS. (2006) 'Spatiotemporal stability of two beetle populations in non-farmed habitats in an agricultural landscape' Journal of Applied Ecology 43 (4), pp 680 - 689

Blackshaw, RP. & Goodwin, T. (2004) 'Complex patterns from simple decisions: the emergence of behaviours in Otiorhynchus sulcatus F' Agricultural and Forest Entomology 6 (4), pp 285 - 290

Petrovskii, SV. & Blackshaw, RP. (2003) 'Behaviourally structured populations persist longer under harsh environmental conditions' Ecology Letters 6 (5), pp 455 - 462

Murchie, AK., Moore, JP., KFA, W. & Blackshaw, RP. (2003) 'Invasion of agricultural land by the earthworm predator, Arthurdendyus triangulatus (Dendy)' Pedobiologia 47 (5-6), pp 920 - 923

Blackshaw, RP. & Coll, C. (1999) 'Economically important leatherjackets of grassland and cereals: biology, impact and control' Integrated Pest Management Review 4 (2), pp 145 - 162 http://www.springerlink.com/content/1353-5226/4/2/

McGee, C., Wisdom, GB., Fairweather, I., Blackshaw, RP., McIlroy, J. & Walker, B. (1998) 'Characterization of the proteins present in the mucus of the flatworm Artioposthia triangulata (Dendy)' COMP BIOCHEM PHYS B 119 (2), pp 293 - 298

Haria, AH., McGrath, SP., Moore, JP., Bell, JP. & Blackshaw, RP. (1998) 'Impact of the New Zealand flatworm (Artioposthia triangulata) on soil structure and hydrology in the UK.' Sci Total Environ 215 (3), pp 259 - 265

Blackshaw, RP. & DArcyBurt, S. (1997) 'Spatial distribution of bibionid larvae in agricultural grassland' ENTOMOL EXP APPL 84 (1), pp 17 - 25

McGee, C., Fairweather, I. & Blackshaw, RP. (1997) 'Ultrastructural features of the epidermis of the planarian Artioposthia triangulata (Dendy)' HYDROBIOLOGIA 347, pp 15 - 24

Blackshaw, RP., Moore, JP. & Alston, R. (1996) 'Removal trapping to control Artioposthia triangulata' ANN APPL BIOL 129 (2), pp 355 - 360

McGee, C., Fairweather, I. & Blackshaw, RP. (1996) 'Ultrastructural observations on rhabdite formation in the planarian, Artioposthia triangulata' J ZOOL 240, pp 563 - 572

Blackshaw, RP. (1995) 'Changes in populations of the predatory flatworm Artioposthia traingulata and its earthworm prey in grassland.' Acta Zoologica Fennnica 196, pp 107 - 110 http://www.sekj.org/ActaZool.html

BLACKSHAW, RP. & PERRY, JN. (1994) 'PREDICTING LEATHERJACKET POPULATION FREQUENCIES IN NORTHERN-IRELAND' ANN APPL BIOL 124 (2), pp 213 - 219

BLACKSHAW, RP. & DARCYBURT, S. (1993) 'A COMPARISON OF SAMPLING METHODS FOR BIBIONID LARVAE (DIPTERA, BIBIONIDAE) IN GRASSLAND' ANN APPL BIOL 123 (3), pp 527 - 530

BLACKSHAW, RP. & THOMPSON, D. (1993) 'COMPARATIVE EFFECTS OF BARK AND PEAT BASED COMPOSTS ON THE OCCURRENCE OF VINE WEEVIL LARVAE AND THE GROWTH OF CONTAINERIZED POLYANTHUS' J HORTIC SCI 68 (5), pp 725 - 729

HUMPHREYS, IC., BLACKSHAW, RP., STEWART, RM. & COLL, C. (1993) 'DIFFERENTIATION BETWEEN LARVAE OF TIPULA-PALUDOSA AND TIPULA-OLERACEA (DIPTERA, TIPULIDAE) USING ISOELECTRIC-FOCUSING, AND THEIR OCCURRENCE IN GRASSLAND IN NORTHERN BRITAIN' ANN APPL BIOL 122 (1), pp 1 - 8

STEWART, VI. & BLACKSHAW, RP. (1993) 'GENETIC-VARIATION IN POPULATIONS OF THE TERRESTRIAL PLANARIAN ARTIOPOSTHIA-TRIANGULATA (DENDY), AND EVIDENCE FOR PASSIVE DISPERSAL IN NORTHERN-IRELAND' ANN APPL BIOL 123 (2), pp 459 - 468

Blackshaw, RP. & Stewart, VI. (1992) 'Artioposthia triangulata (Dendy, 1894), a predatory terrestrial planarian and its potential impact on lumbricid earthworms.' Agricultural Zoology Reviews 5, pp 201 - 219

BLACKSHAW, RP. (1992) 'BLACK VINE WEEVIL (OTIORHYNCHUS-SULCATUS (F)) OVIPOSITION ON POLYANTHUS PLANTS OUTDOORS IN NORTHERN-IRELAND' J HORTIC SCI 67 (5), pp 641 - 646

Blackshaw, RP. (1992) 'Flatworm studies' New Farmer and Grower 34, p 36

BLACKSHAW, RP. (1992) 'THE EFFECT OF STARVATION ON SIZE AND SURVIVAL OF THE TERRESTRIAL PLANARIAN ARTIOPOSTHIA-TRIANGULATA (DENDY) (TRICLADIDA, TERRICOLA)' ANN APPL BIOL 120 (3), pp 573 - 578

BLACKSHAW, RP. & DARCYBURT, S. (1992) 'THE GROWTH OF BIBIO-JOHANNIS (L) AND DILOPHUS-FEBRILIS (L) (DIPTERA, BIBIONIDAE) LARVAE IN THE FIELD' ANN APPL BIOL 120 (2), pp 329 - 337

Blackshaw, RP. (1992) 'The worm in the heart?' Technology Ireland 23 (8), pp 38 - 40

Blackshaw, RP. (1992) 'Threat to our earthworms?' The Biologist 39 (1), pp 4 - 5

DARCYBURT, S. & BLACKSHAW, RP. (1991) 'BIBIONIDS (DIPTERA, BIBIONIDAE) IN AGRICULTURAL LAND - A REVIEW OF DAMAGE, BENEFITS, NATURAL ENEMIES AND CONTROL' ANN APPL BIOL 118 (3), pp 695 - 708

Blackshaw, RP. (1991) 'Killer flatworm' Henry Doubleday Research Association Newsletter 125, pp 32 - 33

BLACKSHAW, RP. (1991) 'MORTALITY OF THE EARTHWORM EISENIA-FOETIDA (SAVIGNY) PRESENTED TO THE TERRESTRIAL PLANARIAN ARTIOPOSTHIA-TRIANGULATA (DENDY) (TRICLADIDA, TERRICOLA)' ANN APPL BIOL 118 (3), pp 689 - 694

BLACKSHAW, RP. & SENTHAMIZHSELVAN, M. (1991) 'THE EFFECT OF SAND PARTICLE-SIZE ON STEINERNEMA-FELTIAE SENSU FILIPJEV (1934) (= N-BIBIONIS BOVIEN) ACTIVITY AGAINST GALLERIA-MELLONELLA LARVAE' ANN APPL BIOL 118 (3), pp 637 - 643

Blackshaw, RP. (1991) 'The New Zealand flatworm' White Admiral: the Newsletter of the Suffolk Naturalists Trust 18, pp 20 - 21 http://www.boxvalley.co.uk/nature/sns/wad.asp

BLACKSHAW, RP. (1990) 'APPLICATION OF A SEQUENTIAL SAMPLING TECHNIQUE TO DIY ASSESSMENT OF LEATHERJACKET (DIPTERA, TIPULIDAE) POPULATIONS IN GRASSLAND' GRASS FORAGE SCI 45 (3), pp 257 - 262

BLACKSHAW, RP. (1990) 'OBSERVATIONS ON THE DISTRIBUTION OF LEATHERJACKETS IN NORTHERN-IRELAND' ANN APPL BIOL 116 (1), pp 21 - 26

BLACKSHAW, RP. (1990) 'STUDIES ON ARTIOPOSTHIA-TRIANGULATA (DENDY) (TRICLADIDA, TERRICOLA), A PREDATOR OF EARTHWORMS' ANN APPL BIOL 116 (1), pp 169 - 176

SENTHAMIZHSELVAN, M. & BLACKSHAW, RP. (1990) 'THE INFLUENCE OF NEOAPLECTANA-BIBIONIS AND HETERORHABDITIS-HELIOTHIDIS AND THEIR ASSOCIATED BACTERIA ON OXYGEN-CONSUMPTION IN GALLERIA-MELLONELLA' J INVERTEBR PATHOL 56 (1), pp 20 - 24

BLACKSHAW, RP. (1989) 'THE EFFECTS OF A CALCAREOUS SEAWEED PRODUCT ON EARTHWORMS IN GRASSLAND SOIL' BIOL AGRIC HORTIC 6 (1), pp 27 - 33

BLACKSHAW, RP. (1988) 'A SURVEY OF INSECT PARASITIC NEMATODES IN NORTHERN-IRELAND' ANN APPL BIOL 113 (3), pp 561 - 565

BLACKSHAW, RP. (1988) 'AN INVESTIGATION INTO THE EFFICACY OF CHLORPYRIFOS AGAINST BIBIONID AND LEATHERJACKET LARVAE IN GRASSLAND' ANN APPL BIOL 112, pp 10 - 11

BLACKSHAW, RP. (1988) 'EFFECTS OF CULTIVATIONS AND PREVIOUS CROPPING ON LEATHERJACKET POPULATIONS IN SPRING BARLEY' RES DEV AGRIC 5 (1), pp 35 - 37

BLACKSHAW, RP. & ONEILL, S. (1987) 'CHLORPYRIFOS COMPOST TREATMENT FOR VINE WEEVIL CONTROL - GROWTH OF ORNAMENTAL PLANTS COMPARED WITH AN ALDRIN STANDARD' J HORTIC SCI 62 (1), pp 67 - 69

DARCYBURT, S. & BLACKSHAW, RP. (1987) 'EFFECTS OF TRAP DESIGN ON CATCHES OF GRASSLAND BIBIONIDAE (DIPTERA, NEMATOCERA)' B ENTOMOL RES 77 (2), pp 309 - 315

BLACKSHAW, RP. & NEWBOLD, JW. (1987) 'INTERACTIONS BETWEEN FERTILIZER USE AND LEATHERJACKET CONTROL IN GRASSLAND' GRASS FORAGE SCI 42 (4), pp 343 - 346

BLACKSHAW, RP. & NEWELL, CR. (1987) 'STUDIES ON TEMPERATURE LIMITATIONS TO HETERORHABDITIS-HELIOTHIDIS ACTIVITY' NEMATOLOGICA 33 (2), pp 180 - 185

BLACKSHAW, RP. (1987) 'STUDIES ON THE CHEMICAL CONTROL OF VINE WEEVIL LARVAE IN HARDY-ORNAMENTAL NURSERY STOCK' J HORTIC SCI 62 (1), pp 63 - 66

BLACKSHAW, RP. (1987) 'STUDIES ON THE ESTIMATION OF SAMPLING ERROR FOR LEATHERJACKETS (TIPULA SPP) IN GRASSLAND' GRASS FORAGE SCI 42 (4), pp 347 - 351

BLACKSHAW, RP. (1987) 'THE USE OF WATER TRAPS FOR CRANE-FLY MONITORING - AN ASSESSMENT OF OPERATOR EFFICIENCY' ANN APPL BIOL 110 (2), pp 239 - 245

BLACKSHAW, RP. (1986) 'AN EVALUATION OF 7 CONTROLLED-RELEASE INSECTICIDES FOR THE CONTROL OF VINE WEEVIL LARVAE' J HORTIC SCI 61 (1), pp 109 - 111

Blackshaw, RP. (1986) 'Leatherjackets: a major grassland pest' Agriculture in Northern Ireland 1 (1), p 20

Moore, JP. & Blackshaw, RP. (1986) 'Protect your stored grain' Agriculture in Northern Ireland 61, pp 132 - 134

BLACKSHAW, RP. (1986) 'RESOLVING ECONOMIC DECISIONS FOR THE SIMULTANEOUS CONTROL OF 2 PESTS, DISEASES OR WEEDS' CROP PROT 5 (2), pp 93 - 99

BLACKSHAW, RP. (1985) 'A PRELIMINARY COMPARISON OF SOME MANAGEMENT OPTIONS FOR REDUCING GRASS LOSSES CAUSED BY LEATHERJACKETS IN NORTHERN-IRELAND' ANN APPL BIOL 107 (2), pp 279 - 285

Mercer, PC. & Blackshaw, RP. (1985) 'Cereal diseases and pests 1984/85' Agriculture in Northern Ireland 60, pp 309 - 312

BLACKSHAW, RP. (1985) 'TESTS OF CONTROLLED-RELEASE INSECTICIDES FOR THE CONTROL OF VINE WEEVIL LARVAE' ANN APPL BIOL 106, pp 16 - 17

BLACKSHAW, RP. (1984) 'AN EVALUATION OF 3 INSECTICIDES AS COMPOST TREATMENTS FOR THE CONTROL OF VINE WEEVIL LARVAE IN HARDY-ANNUAL NURSERY STOCK' J HORTIC SCI 59 (4), pp 559 - 562

Blackshaw, RP. & Mercer, PC. (1984) 'Cereal diseases and pests in 1983/84' Agriculture in Northern Ireland 59, pp 265 - 268

Blackshaw, RP. & Moore, JP. (1984) 'Morphological determination of leatherjacket (Tipula paludosa) instars (Diptera: Tipulidae).' Record of Agricultural Research - Department of Agriculture for Northern Ireland 32, pp 87 - 89

BLACKSHAW, RP. (1984) 'THE IMPACT OF LOW NUMBERS OF LEATHERJACKETS ON GRASS YIELD' GRASS FORAGE SCI 39 (4), pp 339 - 343

Mercer, PC. & Blackshaw, RP. (1983) 'Cereal pests and diseases in 1982/83' Agriculture in Northern Ireland 58, pp 255 - 258

Blackshaw, RP. (1983) 'Earthworms and agriculture' Agriculture in Northern Ireland 58, pp 53 - 55

Blackshaw, RP. & Moore, JP. (1983) 'Hedgerow damage by small ermine moth' Agriculture in Northern Ireland 58, pp 275 - 276

BLACKSHAW, RP. (1983) 'SOME FACTORS INFLUENCING THE ECONOMICS OF PESTICIDE USE IN GRASSLAND' CROP PROT 2 (4), pp 447 - 454

BLACKSHAW, RP. (1983) 'SOME FACTORS INFLUENCING VARIABILITY IN WATER-TRAP CATCHES OF TIPULA SPP (DIPTERA, TIPULIDAE)' B ENTOMOL RES 73 (4), pp 693 - 699

BLACKSHAW, RP. (1983) 'TESTS OF INSECTICIDES AGAINST VINE WEEVIL LARVAE' ANN APPL BIOL 102, pp 36 - 37

BLACKSHAW, RP. (1983) 'THE ANNUAL LEATHERJACKET SURVEY IN NORTHERN-IRELAND, 1965-82, AND SOME FACTORS AFFECTING POPULATIONS' PLANT PATHOL 32 (3), pp 345 - 349

Blackshaw, RP. (1981) 'The Collembola of Northumberland and Durham: an historical record.' Vasculum 66 (not known (4pa)), pp 28 - 37 http://www.the-vasculum.com/

Benefer, CM., Knight, ME., Ellis, JS., Hicks, H. & Blackshaw, RP. 'Understanding the relationship between adult and larval Agriotes distributions: The effect of sampling method, species identification and abiotic variables' Applied Soil Ecology 53, pp 39 - 48 , DOI: 10.1016/j.apsoil.2011.11.004

Letters

BLACKSHAW, R. (1989) 'WORMS OUT OF TURN' NEW SCI 124 (1685), pp 73 - 74

Additional information

External Voluntary Work:

Chair, Dartmoor National Park Authority Standards Committee